Indian Journal of Science and Technology

Nishimiya O. ^1* ,Kunihiro Y. ¹, HiramatsuN. ^1,2, InagawaH.³ ,Todo T. ¹, Matsubara T.⁴ , Reading B.J.² , Sullivan C.V.² and Hara A. ¹

¹ Graduate School of Fisheries Sciences, Hokkaido University, Minato 3-1-1, Hakodate, Hokkaido, 041-8611, Japan. Fax: +81-138-40-8878 email:  [email protected]
² Department of Biology, North Carolina State University, Raleigh, NC 27695-7617, USA
³ Department of Applied Aquabiology, National Fisheries University, Shimonoseki, Yamaguchi 759-6595, Japan.
⁴ South Ehime Fisheries Research Center, Ehime University, Funakoshi 1289-12, Ainan-cho, Minami Uwa, Ehime 798-4292, Japan